Linguistic is a mature science and we should not expect any groundbreaking findings from a new paper. What can do a modern linguist is to have a new interpretation of already known facts, or at best he can find a few new lexical parallels. Nielsen's paper is important because it shows that the linguistic data do not contradict to the available genetic data about the origins of Armenians. And more important it can reinforce it, giving new details.
Creanza et al. 2015 analyzed more than 2000 languages and based that came to the conclusion that the phonetic systems of two neighboring languages correlates better with geographic distance than with their affiliated linguistic families. Another interesting conclusion was that an isolated language drifts. But unlike the genetics were the drift decrease the diversity, the drift in the language increases the number of phonemes.
This by the way can explain the high number of phonemes in the North Caucasian languages. Returning to the Armenian and Kartvelian we can say that similar phonetic systems mean relatively long period of coexistence.
On the other hand, this do not mean that the Kur-Araxian culture which became the substrate for the Armenian language was necessarily and predominantly Kartvelian. First the number of mutual loanwords is too small for that. Most loanwords in Armenian are from the Zan branch which can be connected to the Colchian archaeological culture. Moreover, we do not know well about the phonetic system of Urartian. Given that we know about their phonology only via the cuneiform it is possible that their phonetic system was also close to the Armenian.
Another important question is the migration of IE Anatolians. If further genetic data do not show evidence that they migrated via Balkans, then the only available option left will be their origins or migration via the historic Armenia. In this scenario Kura-Araxes becomes indispensable for their spread. So, there is a need to look at the possible IE Anatolian substrate in Armenian. Nielsen mentions this.
And finally based on the available genetic and archaeological data there is little doubt that the Nakh-Daghestani linguistic family descend from a subset of Kura-Araxes. We can't now deduce the exact boundary of this subset, but we can expect that Proto-Armenian would have a contact with them also. So, another analyze of Armenian and Nakh-Daghestani connections is needed.
Returning to the Kartvelian family. Based on the current genetic and archaeological data the most likely cultures related to the Kartvelian family were the Proto-Colchian (2700-1700/1500BCE) and the Colchian culture (1700/1500-700BCE) in the western Georgia. Some Kartvelian presence in Koban culture is also possible but it's not relevant for ancient Armenian connections. What is more important is that Colchian culture axes were found in northwestern regions of historic Armenia which can explain the stronger presence of Zan loanwords in the Armenian. More ancient DNA and archaeological research from that region, will help to better understand this question.
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