Friday, April 26, 2024

The Yamnaya culture and it's legacy

Yamnaya was an Early Bronze Age culture (3300-2700BC) that expanded in Pontic Caspian steppe in East Europe. It is remarkable due to its homogenous population which contrasts to the diversity of the preceding Eneolithic (means Chalcolithic) cultures. In just 1000 years genes associated with Yamnaya culture reached what is now Portugal and western Mongolia, Britain and South-Central Asia. The scale of this expansion can be considered as one of the most important demographic events in West Eurasia after the invention of farming in Fertile Crescent. In 2022 Lazaridis et al. announced that all modern living IE nations languages can be derived from the Yamnaya.
In 2015 the first ancient DNA of Yamnayans was published and it became clear that they do not directly descend from Eastern European hunter gatherers. Roughly the half of their ancestry was from south, and the "Armenian like" term was used to label this southern part. Then in late 2015 the Caucasian hunter gatherers (CHG) were discovered. In 2016 Iranian Zagros Neolithic farmers were published and since then both those populations were used to model the Yamnayans.
Despite those statistical models the formation of Yamnayans gene pool was remaining an obscure subject. Finally in April 2024 two large papers dedicated to this subject provided more clarity. Various Neolithic and Eneolithic groups living in East Europe had both shared ancestry and differences. One group who was living from North Caucasus foothills to Lower Volga (site Berezhnovka ) was labeled as "Caucasus Lower Volga cline". CLV had both southern Caucasian hunters and farmers (Aknashen) ancestry.
I will dedicate a special topic to describe what is CLV but at this stage it's sufficient to say that Yamnayans had up to 80% of their ancestry from CLV. Not only Yamnayans but all Kurganic cultures (Sredni Stog, Khvalynsk, Cernabida, Usatovo) of eastern Europe and Balkans had the CLV ancestry mixed with various local European populations. For the case of Yamnayans the second part of their ancestry was the Ukraine hunter gatherers (UNHG), which made them similar to Sredni Stog. Even though the most important Y DNA's were missing in Sredni Stog which could mean that Proto Yamnaya's exact immediate geographic location is not found yet but shouldn't be outside from Don - Volga - Caucasus triangle. With more sampling we will learn the exact origin of the two most successful lineages. R1b-M269 and R1a-M417.
The samples from the new paper are not available yet but we have two samples from the forementioned CLV cline. They are from the Progress - 2 site in the North Caucasus (4500-4000BC, see the last map for the location). I used them to model various Yamnaya samples and got the same result as in Lazaridis 2024. More than 75-80% of ancestry in regular Yamnaya samples is from the Progress / CLV. Some outliers have extra ancestries which is normal given their location.
You may wonder how "Hittites from east, from historic Armenia" and "Yamnaya from CLV" are connected to each other? After all there is no evidence that Hittites and other Anatolian speakers descend from Yamnaya culture. What is the common shared ancestry between IE Anatolians and Yamnaya?
Another topic will be dedicated to this subject.
And at the end here is the list of Y DNA found in Yamnaya at this stage. R1b-Z2103 (west Asia Balkans), R1b-L51 (west Europe), R1a-M417 (Asia - Europe), I2-L699, J2b-L283 (west Balkans), R1b-PF7562 (Balkans Mycenean ) Q-L939 and isolated cases of J1 and R1b-V1636.

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